2015 - Current

41. S. Lobionda, S. Hur, J. Jeong, Y.K. Lee#, and Y. Hwang#: "Synthetic matrix-mediated cell spreading and its roles in M1 macrophage polarization through a Rho/ROCK signaling pathway" (in preparation) (#denotes co-corresponding authors)

40. K. Ahn, E. Park, C. Choi, H. Cha, Y. Hwang# and S. Nam#: “Hyaluronic Acid Treatment Improves Healing of the Tenorrhaphy Site by Suppressing Adhesion through Extracellular Matrix Remodeling in a Rat Model” Polymers, 2021, 13(6), 928 (Link) (#denotes co-corresponding authors) 

39.  J. Kim, S. Park, H. Choi, Y. Hwang, M. Ban, and J. Park: “Cotton ball self-retraction technique: a novel surgical method used during microflap dissection” Laryngoscope, 2021. (in press) (Link)

38. H. Kim, J. Jeong, M. Fendereski, H. Lee, D. Kang, Y. Kim, N. Pham, S. Hur, J. Amirian, N. Suh, N. Hwang, S. Ryu, J. Yoon#, and Y. Hwang:#: "Heparin-mimicking polymer-based in vitro platform to recapitulate in vivo muscle atrophy phenotypes" International Journal of Molecular Sciences, 2021, 22(5), 2488. (Link) (#denotes co-corresponding authors)

37. H. Kim, L. Anggradita, S. Lee, S. Hur, N. S. Hwang, S. Nam#, and Y. Hwang#: “Ameliorating fibrotic phenotypes of keloid skin-derived dermal fibroblasts through a EGF signaling pathway” International Journal of Molecular Sciences, 2021, 22(4), 2198. (Link) (#denotes co-corresponding authors)

36. S. Hong, S. Oh, D. Choi, Y. Hwang#, and S. Kim#: "Self-organized Liver Microtissue on a Bio-functional Surface: the Role of Human Adipose-derived Stromal Cells in Hepatic Function", International Journal of Molecular Sciences, 2020, 21(13), 4605. (#denotes co-corresponding authors) (Link)

35. S. Kim, J. Nam, M. Lee, Y. Hwang, and E. Park: "Effects on wound healing of human induced pluripotent stem cell-derived cells similar to endothelial colony-forming cells", Journal of Wound Management and Research, 2020, 16(1), 3-12. (Link)

34. S. Hur, J Jeong, J Yoon, and Y. Hwang: "Traction force microscopy for understanding cellular mechanotransduction", BMB Reports, 2020, 53(2), 74-81. (Link)

33. M. Hepburn, P. Wijesinghe, L, Major, J. Li, A. Mowla, C. Astell, H. Park, Y. Hwang, Y. Choi, and B. Kennedy: "Three-dimensional imaging of cell and extracellular matrix elasticity using quantitative micro-elastography", Biomedical Optics Express, 2020, 11(2), 867-884. (Link)

32. C. Kim, J. Young, A. Holle, K. Jeong, L. Major, J. Jeong, Z. Aman, D. Han, Y. Hwang, J. Spatz, Y. Choi: "Stem cell mechanosensation on gelatin methacrylol (GelMA) stiffness gradient hydrogels", Annals of Biomedical Engineering, 2020, 48, 893-902. (Link)

31. L. Major, A. Holle, J. Young, M. Hepburn, K. Jeong, I. Chin, R. Sanderson, J. Jeong, Z. Aman, B. Kennedy,  D. Han, Y. Hwang, H. Park, K. Guan, J. Spatz, Y. Choi: "Volume adaptation controls stem cell mechanotransduction", ACS Applied Materials and Interfaces, 2019, 11, 45520-45530 (Link).

30. D. Han, C. Ahn, J. Lee, Y. Hwang, J. Kim, K. Park, J. Lee, and K. Son: "Optimization of electrospun poly(caprolactone) fiber diameter for vascular scaffolds to maximize smooth muscle cell infiltration and phenotype modulation", Polymers, 2019, 11(4), 643 (Link).

29.  J. Sah, N. Hao, Y. Kim, T. Eigler, E. Tzahor, S. Kim#, Y. Hwang#, and J. Yoon#: “MBP-FGF2-immobilized matrix maintains the self-renewal and myogenic differentiation potential of skeletal muscle stem cells”, International Journal of Stem Cells, 2019, 12(2), 360-366. (#denotes co-corresponding authors) (Link)

28. H. Kim, Y. Kim, J. Park, N.S. Hwang, Y. Lee, and Y. Hwang: "Recent Advances in Engineered Stem Cell-derived Cell Sheets for Tissue Regeneration", Polymers, 2019, 11(2), 209. (Link)

27. T. Ha, H. Kang, T, Kim, J. Kwon, H. Kim, A. Ryu, H. Jeon, J. Han, H. Broxmeyer, Y. Hwang#, Y. Lee#, and M. Lee#: “MiR-9 controls chemotactic activity of cord blood CD34+ cells by repressing CXCR4 expression” International Journal of Stem Cell, 2018, 11(2), 187-195. (#denotes co-corresponding authors) (Link)

26. H. Kim, Y. Kim, M. Fendereski, N. S. Hwang, and Y. Hwang: "Recent Advancements in Decellularized Matrix-based Biomaterials for Musculoskeletal Tissue Regeneration" Adv Exp Med Biol: Novel Biomaterials for Regenerative Medicine, 2018, 1077, 149-162. (Link)


25. C. Ahn, Y. Kim, S. Park, Y. Hwang#, and J. Lee#: "Development of arginine-glycine-aspartate-immobilized 3D printed poly(propylene fumarate) scaffolds for cartilage tissue engineering" Journal of Biomaterials Science, Polymer Ed., 2018, 29(7-9), 917-931. (# denotes co-corresponding) (Link)

24. H. Kim, Y. Lee, Y. Kim, Y. Hwang#, and N. Hwang#: “Biomimetically reinforced polyvinyl alcohol-based hybrid scaffolds for cartilage tissue engineering” (#denotes co-corresponding authors) Polymers, 2017, 9(12), 655. (Link)

23. Y. Hwang, T. Seo, S. Hariri, C. Choi, and S. Varghese: “Matrix topographical cue-mediated myogenic differentiation of human embryonic stem cell derivatives” Polymers, 2017, 9(11), 580. (Link)

22. Y. Hwang, H. Broxymeyer, and M. Lee: “Generating autologous hematopoietic cells from human-induced pluripotent stem cells through ectopic expression of transcription factors” Curr Opin Hematol, 2017, 24(4), 283-88.60/9/1/13. (Link)

21.  Y. Hwang, S. Park, and J. Lee: “Applications of Functionalized Carbon Nanotubes for the Therapy and Diagnosis of Cancer” Polymers, 2017, 9(1), 13. (Link)

20. C. Wen, H. Kang, Y. Shih, Y. Hwang, and S. Varghese:"In vivo comparison of biomineralized scaffold-directed osteogenic differentiation of human embryonic and mesenchymal stem cells" Durg Delivery and Translational Research, 2016, 6(2), 121-31. (Link)


19. M. Nakasaki, Y. Hwang, Y. Xie, S. Kataria, R. Gund, E. Y. Hajam, R. Samuel, R. George, D. Danda, P. MJ, T. Nakamura, Z. Shen, S. Briggs, S. Varghese, and C. Jamora: “The matrix protein Fibulin-5 lies at the interface of tissue stiffness and inflammation in fibrosis” Nature Communications, 2015, 6:8574. (Link)


18. H. Kabra#, Y. Hwang#, M. Kar, and S. Varghese: “Biomimetic material-based delivery of stem cells enhances in vivo engraftment” ACS Biomaterials Science and Engineering, 2015, 1, 7-12. (# denotes joint first author) (Link)


17. H. Kim, J. Heo, Y. Hwang, S. Kwak, O. Park, H. Kim, S. Varghese, and N. Hwang: “Extracellular matrix-based Arg-Gly-Asp modified photopolymerizing hydrogels for cartilage tissue engineering” Tissue Engineering Part A, 2015, 21(3-4), 757-766. (Link)

2009 - 2014

16. H. Kang, Y-R. V. Shih, Y. Hwang, C. Wen, T. Seo, and S. Varghese: “Mineralized gelatin methacrylate-based matrices induce osteogenic differentiation of human induced pluripotent stem cells” Acta Biomaterialia, 2014, 10(12), 4961-4970. (Link)


15. H. Kang, C. Wen, Y. Hwang, Y. Shih, M. Kar, S. Seo, and S. Varghese: “Biomaterials matrix-assisted osteogenic differentiation of human embryonic stem cells” Journal of Materials Chemistry B, 2014, 2(34), 5676-5688. (Link)


14. Y. Hwang, S. Suk, T. Seo, B. Du, Y. Xie, Z. Li, and S. Varghese: “WNT3A promotes myogenesis of human embryonic stem cells and enhances in vivo engraftment” Scientific Reports, 2014, 4, 5916; DOI: 10.1038/srep05916. (Link)


13. H. Lim, Y. Hwang, M. Kar, and S. Varghese: “Smart hydrogels as functional biomimetic systems” Biomaterials Science, 2014, 2, 603-618. (Link)


12. Y-R. V. Shih, Y. Hwang, A. Phadke, H. Kang, N. Hwang, E. Caro, S. Nguyen, M. Siu, E. Theodorakis, N. Gianneschi, K. Vecchio, S. Chien, O. Lee, and S. Varghese: “Calcium phosphate-bearing matrices induce osteogenic differentiation of stem cells through adenosine signaling.” Proceedings of National Academy of Science, 2014, 111(3): 990-5. (Link)

  • Featured in UCSD news (January 6, 2014)

  • Featured in CIRM news (January 7, 2014)

  • Selected as Top story in Extracellular Matrix News 5.00 (January 9, 2014) 



11. Y. Hwang#, S. Suk#, M. Tierney, S. Lin, B. Du, T. Seo, A. Mitchell, A. Sacco, and S. Varghese: “Directed in vitro myogenesis of human embryonic stem cells and their in vivo engraftment” PLoS One, 2013, 8(8): e72023. (# denotes joint first author) (Link)


10. A. Phadke#, Y. Hwang#, S. Kim, S. Kim, T. Yamaguchi, K. Masuda, and S. Varghese: “Effect of scaffold microarchitecture on osteogenic differentiation of human mesenchymal stem cells” European Cells & Materials, 2013, 25, 114-29. (# denotes joint first author) (Link)


9. C. Chang#, Y. Hwang#, D. Brafman#, T. Hagan, C. Phung, and S. Varghese: “Engineering cell-material interfaces for long-term expansion of human pluripotent stem cells” Biomaterials, 2013, 34(4), 912-21. (# denotes joint first author) (Link)

  • Selected as Top story in Extracellular Matrix News 3.44, November 8, 2012



8. Q. Nguyen, Y. Hwang, A. Chen, S. Varghese, and R. Sah: “Cartilage-like mechanical properties of poly(ethylene glycol)-diacrylate hydrogels” Biomaterials, 2012, 33(28), 6682-90. (Link)



7. C. Zhang, N. Sangaj, Y. Hwang, A. Phadke, C. Chang, and S. Varghese: “Oligo(trimethylene carbonate)-poly(ethylene glycol)- oligo(trimethylene carbonate) triblock based hydrogels for cartilage tissue engineering” Acta Biomaterialia, 2011, 7(9), 3362-3369. (Link)


6. Y. Hwang#, A. Phadke#, and S. Varghese: “Engineered microenvironments for self-renewal and musculoskeletal differentiation of stem cells” Regenerative Medicine, 2011, 6(4), 505-24. (# denotes joint first author) (Link)


5. R. Ayala, C. Zhang, D. Yang, Y. Hwang, A. Aung, S. Shroff, F. Arce, R. Lal, G. Arya, and S. Varghese: “Engineering the cell-material interface for controlling stem cell adhesion, migration, and differentiation” Biomaterials, 2011, 32(15), 3700-11. (Link)

4. A. Phadke, C. Zhang, Y. Hwang, K. Vecchio, and S. Varghese: “Templated mineralization of synthetic hydrogels for bone-like composite materials: Role of matrix hydrophobicity” Biomacromolecules, 2010, 11(8), 2060–68. (Link)


3. Y. Hwang, N. Sangaj, and S. Varghese: “Interconnected macroporous poly (ethylene glycol) (PEG) cryogels as a cell scaffold for cartilage tissue engineering” Tissue Engineering Part A, 2010, 16(10), 3033-41. (Link)


2. Y. Hwang, C. Zhang, and S. Varghese: “Poly (ethylene glycol) cryogels as potential cell scaffolds: Effect of polymerization conditions on cryogel microstructure and properties” Journal of Materials Chemistry, 2010, 20(2), 345-51. (Link)

  • Selected as Advance Articles for Highlights in Chemical Biology

1. N. Hwang, C. Zhang, Y. Hwang, S. Varghese: “Mesenchymal stem cell differentiation and roles in regenerative medicine” Wiley Interdisciplinary Reviews: Systems Biology and Medicine, 2009, 1(1), 97-106. (Link)